Revised Pages Key to the Genera of Leptosporangiate and

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Michigan Ferns and Lycophytes,A Guide to Species of the Great Lakes Region. Daniel D Palmer, https www press umich edu 9742700 michigan ferns and lycophytes. 66 MICHIGAN FERNS AND LYCOPHY TES, tips of fronds or sterile fronds and upper and lower divisions of fertile. frond 1 pinnate pinnatifid with much smaller darker fertile pinnae be. tween fertile parts not persisting through winter Osmunda. 7 Sterile blades pinnatifid to 1 pinnate pinnatifid sterile and fertile pinnae. arising separately from ground fertile blades shorter hard dark and per. sisting through winter 8, 8 7 Sterile blades 1 pinnate pinnatid throughout rachises not winged veins. free Matteuccia, 8 Sterile blades pinnatifid rachises winged the lowest pinnae sometimes.
free veins netted Onoclea, 9 6 Sori linear or slightly curved along veins or parallel to costae or in contin. uous bands along margins of pinnae under a false indusium 10. 9 Sori round or nearly so along veins from costae toward margin or near. margins of pinnae or pinnules 17, 10 6 Sori long linear along veins cross section of stipe bases reveals 2 back to. back C shaped vascular bundles uniting to form an X shape higher up. stipe scales clathrate Asplenium in part A platyneuron. A ruta muraria A trichomanes A viride, 10 Sori short or long linear along veins pinna or pinnule margins or along. costae of pinna or pinnule midribs cross section at stipe bases reveals 2. straplike parallel vascular bundles scales nonclathrate 11. 11 10 Sori linear or j shaped along veins that extend from midribs toward mar. 11 Sori linear along margins of pinnules or along areolar veins near and par. allel to midrib 14 Asplenium Athyrium Cryptogramma Deparia. Homalosorus Pellaea Pteridium, 12 11 Blades 1 pinnate pinna margins entire fertile fronds narrower and with. much narrower pinnae Homalosorus H pycnocarpos, 12 Blades 1 pinnate pinnatifid to 2 pinnate pinnatisect sterile and fertile.
fronds the same shape or nearly so 13, 13 12 Grooves of pinna costae shallow not continuous with grooves of rachises. blades 1 pinnate pinnatifid sori linear Deparia D acrostichoides. 13 Grooves of pinna costae deep continuous with grooves of rachises blades. pinnate pinnatisect to 2 pinnate pinnatifid sori linear or J shaped. hooked over vein tips Athyrium A filix femina, 14 11 Sori short linear in single chainlike rows on anastomosing areolar veins. on each side of and parallel to costae and sometimes along midribs veins. joining to form a single row of areoles near and on both sides of midribs. then free and branching to margins Woodwardia, 14 Sori short to long linear along pinna or pinnule margins veins free not. forming areolae near costae 15, 15 14 Fronds 35 180 cm long 20 50 cm wide well separated growing in large. colonies on long creeping underground rhizomes sterile and fertile. fronds the same blades broadly triangular growing in loose soil in open. areas stipes straw colored to green or brown Pteridium. 15 Sterile fronds 3 50 cm long 2 20 cm wide fronds clustered in small tufts. Michigan Ferns and Lycophytes,A Guide to Species of the Great Lakes Region.
Daniel D Palmer, https www press umich edu 9742700 michigan ferns and lycophytes. Fronds Anatomy A Frond tip apex B Pinnae C Rachis D Basal pinna E Stipe F. Rhizome G Blade H Frond I Blade J Crosier K Pinna L Stipe M Rhizome N Roots. O Pinna stalk P Pinnule Q Costa R Costule S Rachis T V Ultimate segments. Michigan Ferns and Lycophytes,A Guide to Species of the Great Lakes Region. Daniel D Palmer, https www press umich edu 9742700 michigan ferns and lycophytes. Pinna Forms and Margins,Michigan Ferns and Lycophytes. A Guide to Species of the Great Lakes Region,Daniel D Palmer.
https www press umich edu 9742700 michigan ferns and lycophytes. on short ascending to erect rhizomes sterile blades different usually grow. ing on rock stipes dark purple to black or stipes brown at base green or. straw colored distally 16, 16 15 Stipes dark shiny brown to dark purple to black fronds somewhat dimor. phic or not sterile and fertile pinnae linear oblong to oblong lanceolate. 16 Stipes brown at base green or straw colored distally fronds strongly di. morphic fertile fronds obviously longer than the sterile narrow elongate. usually with revolute pinnules sterile pinnules oblong to fan shaped fer. tile pinnules narrow elongate Cryptogramma, 17 9 Blades deeply pinnatifid rhizome long creeping indusia lacking. Polypodium, 17 Fronds 1 pinnate to 3 pinnate rhizomes erect short or long creeping in. dusia present or absent 18, 18 17 Blades 1 pinnate pinna margins sharply toothed with long stiff bristlelike. projections indusia peltate pinnae eared at base sterile and fertile pinnae. same size or fertile pinnae near frond tips much smaller Polystichum. 18 Blades 1 pinnate pinnatifid to 3 pinnate pinnatifid pinna margins with or. without fine teeth indusia of various types sterile and fertile fronds same. size or different fertile pinnae near frond tips not smaller 19. Michigan Ferns and Lycophytes,A Guide to Species of the Great Lakes Region.
Daniel D Palmer, https www press umich edu 9742700 michigan ferns and lycophytes. 70 MICHIGAN FERNS AND LYCOPHY TES,Scales and Hairs. 19 18 Indusia absent fronds ternate or broadly triangular with winged rachises. 19 Indusia of various types present fronds not ternate or broadly triangular. rachises not winged 21, 20 19 Fronds ternate rachises not winged stipes 1 5 to 3 times length of blade. costae grooved adaxially lacking small white sharply pointed acicular. hairs Gymnocarpium, 20 Fronds broadly triangular rachises winged stipes 1 to 1 5 length of blade. costae not grooved adaxially small white sharply pointed acicular hairs. on undersurface of pinnae and rachis Phegopteris, 21 19 Blades 1 pinnate pinnatifid fine white sharp tipped translucent needlike.
acicular hairs present on rachises and costae cross section at base of stipe. shows two slightly curved linear vascular bundles that unite upward to a. U shape Thelypteris, 21 Blades 1 pinnate to 3 pinnate pinnatifid acicular hairs absent cross sec. tion at base of stipe various not with two linear vascular bundles 22. 22 21 Rhizomes ascending to erect indusia above sori round or reniform. 1 pinnate pinnatifid to 3 pinnate pinnatifid glandular or not. Dryopteris,Michigan Ferns and Lycophytes,A Guide to Species of the Great Lakes Region. Daniel D Palmer, https www press umich edu 9742700 michigan ferns and lycophytes. Sori and Indusia, 22 Rhizomes ascending to erect or long creeping subterranean sori with cup. like or tubular indusia or ribbonlike or filamentous hairs under indusion. spreading to cover it 23, 23 22 Rhizomes long creeping below soil surface stipes and rachises hairy.
scales lacking sori marginal at vein tips in sinuses small less than 0 5. mm in diam cup shaped circular or slightly 2 valvate cups fronds 30 90. long rare Jackson County Dennstaedtia, 23 Rhizomes decumbent to erect sori larger than 0 5 mm stipes and rachis. es scaly with or without hairs sori not marginal indusia cup shaped at. tached at base looking outward or composed of many narrow to hairlike. segments attached beneath and encircling sori fronds shorter than 50 cm. 24 23 Indusia hoodlike soon shriveling and absent opening outward attached. at bases and partially overlapping sori old stipe bases not persistent or. sparsely persistent on rhizomes fronds up to 50 cm long Cystopteris bul. bifera often longer Cystopteris, 24 Indusia composed of many narrow to hairlike segments attached beneath. and encircling sori old stipe bases persist on rhizome fronds up to 25 cm. long Woodsia obtusa sometimes longer Woodsia,Michigan Ferns and Lycophytes. A Guide to Species of the Great Lakes Region,Daniel D Palmer. https www press umich edu 9742700 michigan ferns and lycophytes. 72 MICHIGAN FERNS AND LYCOPHY TES,Adiantum Linneaus Pteridaceae.
Maidenhair Ferns, See Pteridaceae in appendix I for a description of the family and a key to the Mich. igan genera, Etymology Greek adiantos unwetted an ancient name alluding to the water. repellent fronds Maidenhair a name used for obscure reasons is applied to ferns. with delicate fronds and slender black stipes see also Asplenium trichomanes. Plants Monomorphic deciduous fronds in loose clusters in clones. Rhizomes Creeping, Fronds Stipes erect with blade horizontal to ground. Stipes Round usually brown to purple or black shiny mostly glabrous except. at very base sometimes sparsely hairy or with fibrils. Blades 2 pinnate to 3 pinnate,Pinnae Elongate tips obtuse. Pinnules Rhomboidal trapezoidal fan shaped or dimidiate sessile to short. stalked never adnate,Veins Free repeatedly branching dichotomously.
Sori Less than 4 mm long short linear marginal along several vein tips. Indusial flaps Consisting of reflexed margins of pinnae rectangular or U. Sporangia Distinctively borne on undersurface of the false indusia. Chromosome x 29 30, Adiantum is a genus of about 200 species widely distributed in tropical and tem. perate areas mostly at low to mid elevations and in wet forests Nine species are. treated in the Flora of North America North of Mexico vol 2 1993 One species. is found in Michigan,Adiantum pedatum Linnaeus,Maidenhair Fern Northern Maid. enhair Fern,Etymology Latin pedatum,footlike the frond outline resembles a. bird s foot Linnaeus used this term to,describe the fronds of this fern. Plants Monomorphic deciduous,Rhizomes Short creeping thick.
often with attached old stipe bases,scales yellowish margins entire. Michigan Ferns and Lycophytes,A Guide to Species of the Great Lakes Region. Daniel D Palmer, https www press umich edu 9742700 michigan ferns and lycophytes. Adiantum pedatum Maidenhair fern A Frond B Pinna detail showing veins and. marginal linear sori, Informative arrows 1 Dimidiate pinnules 2 Marginal linear sori. Michigan Ferns and Lycophytes,A Guide to Species of the Great Lakes Region.
Daniel D Palmer, https www press umich edu 9742700 michigan ferns and lycophytes. 74 MICHIGAN FERNS AND LYCOPHY TES, Stipes 1 2 mm diam erect triangular alternate dimidiate lower. round glabrous dark brown or margin straight acroscopic margins. purplish brown to black stipe tips lobed lobes separated by a narrow. fork to form 2 rachises spreading in a sinuses 1 2 3 mm deep. Y fashion Veins Free forked repeatedly, Fronds 35 75 cm long delicate Sori 1 3 mm long linear to. crosiers reddish brown when young slightly curved marginal on tips of. Blades 15 30 20 40 cm 2 pinate lobes covered with elongate green to. to 3 pinnate semicircular fan gray reflexed marginal false indusia. shaped formed by rachises dividing Habitat Grows in a variety of soils. into 2 branches with several pinnae best in rich mature deciduous shady. forming on the outer edge of the woods in moist alkaline and humus. branches horizontal to ground to covered soils,slightly drooping Distribution Eastern Canada. Pinnae 5 20 cm long elongate USA Northeastern states extending. tips obtuse longest pinnae closest to south to northern Georgia and Okla. stipes homa The distribution is essentially, Pinnules 8 21 pinnule pairs 3 that of the deciduous forests of eastern.
times as long as wide smaller pinnules North America. Some authors divide this species in North America into 3 to 4 subspecies Adi. antum pedatum and A aleuticum which does not occur in Michigan are here. recognized as 2 closely related species in North America. Adiantum pedatum is a graceful fern with distinct fronds that is unique. within the Michigan flora and easily recognized Its silhouette alone is diag. nostic Its horizontal or drooping fan shaped blades are borne on narrow dark. stipes and are divided into 2 equally branched rachises Its forked rachises ra. diate pinna on one side only Its pinnules are on narrow stalks and the linear. sori on their outer edges are attached to the undersurface of an in rolled mar. gin forming a false indusium,ASPLENIUM Linnaeus Aspleniaceae. Spleenworts, Aspleniaceae as recognized here is comprised of a single huge extremely diverse. genus Asplenium Asplenium contains about 700 species worldwide mostly in the. tropics The family and generic treatments are identical and found below. Etymology Greek a not splen In ancient Greece the plants were believed. to cure spleen diseases The vernacular name derives from the Greek spleenworts. Plants Small to medium sized terrestrial or epipetric. Fronds Monomorphic small to medium sized clustered to remote. Revised Pages Ferns 65 Key to the Genera of Leptosporangiate and Eusporangiate Ferns Polypodiopsida as Found in Michigan 1 Plants aquatic or partially so small and free floating or clover shaped and

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